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You can apply to register a mark but another party can object if they are already using the same mark for the same class of goods and services even if their earlier mark isn't a registered mark. They would have existing unregistered rights and you would have immense difficulty in getting your application past the registrar under such circumstances.
Adding .com or .co.uk as a suffix is not relevant to the registrar. It is the substantive element that counts.
There is no direct link between ownership of a domain name and ownership of a similar trademark because there are 45 general classes of goods and services under which a mark can be registered and there is no legal obligation to register a mark before using it to trade. Therefore, even though Megacorp has a registered mark for 'xyz' in 44 classes (so many classes is very unlikely), as long as you are only using xyz.com for goods and services in the one remaining class you are not violating Megacorp's trademark rights. Megacorp doesn't have an automatic right to the registration of the domain. If you try to sell it to Megacorp then you will be exposing yourself to the probability of having the domain taken from you under cybersquatting rules.
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